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August 1, 2025
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mar 16, 1963 - Nikolaas Tinbergen Publishes "On aims and methods of Ethology" (Incorporating "Tinbergen's Four Questions")

Description:

Dutch biologist Niko Tinbergen publishes his paper "On aims and methods of Ethology". This paper outlines a framework for successfully studying animal behavior, including his now famous 4 questions: Causation, Development, Evolution, and Function.

Added to timeline:

Date:

mar 16, 1963
Now
~ 62 years ago